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Retrieved 26 May Among frogs, all of these new communication channels mentioned above, as well as the control and directionality of the vocal sacs such as observed in the present study , and even new integration between distinct communication modes such as visual-tactile bimodal signaling in H. Male communication: Territoriality and signal choice Males defend territories as breeding sites calling and courtship. The solution is a change in the way things are sold in the first place. Without an apparent order, the female performed visual-tactile signals: arm lifting, or arm waving both usually ending with female touching the face of the male or his anterior or posterior dorsal body region, depending on which arm the female uses to perform the signal; Fig 5 and S5 Movie show the position of the couple , or body jerking.

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